We maintain a Zotero bibliographic database of publications from the reserves of the Central Sierra Field Research Stations (CSFRS). Learn more about this list here. See bibliographies for:

Sagehen’s publications include:

Research publications are available from conventional sources like university libraries. Where there is an online version available, we have included the URL in the Zotero bibliographic record. Hard copies of most early Masters and Ph.D. theses produced at Sagehen are available in the Sagehen Creek Field Station library; station collections are for on-site use, only.

Some original Sagehen documents and media are housed in the UC Berkeley Bancroft Library, i.e. station use logs from the 1950’s, A. S. Leopold wildlife course lecture recordings, original data sheets, etc.

See our calendar for abstracts of research and education projects currently underway at Sagehen.

Major partners: UC Natural Reserve System.

Masters & Ph.D. Theses


  1. Ryan, Leslie. Art/sci. 2021? Ph.D., OSU
  2. Jiao, Yi. Natural sources and sinks of BVOCs, especially halocarbons and hydrocarbons. 2021? Ph.D., UCB
  3. Scheckel, Kelsey. Sagehen’s other slave-making ants: Formica aserva. 2021?, Ph.D., UCB
  4. Neuhausler, Rosanna. Investigating critical timescales of water storage and discharge in the Sierra Nevada, California. 2021?, Ph.D., UCB
  5. Natali, Jennifer. Interactions of water, sediment, plants and climate in montane meadow ecosystems of the Sierra Nevada in California. 2021?, Ph.D., UCB


  1. Dietrich, Scott. Habitat, Diet, and Foraging Ecology of Willow Flycatcher in Sierra Nevada Meadows. 2020, Masters, USU
  2. Safa, Hamideh. Water and Energy Budgets of Snow-Covered Montane Forests: Observations, Remote Sensing and Modeling. 2020 Ph.D. UNR
  3. Tubbesing, Carmen. Post-fire tree regeneration in an altered disturbance regime: community dynamics and interactions with land management. 2020, Ph. D., UCB
  4. Klip, Mario. Spatial and temporal patterns of a generalist urban carnivore: American black bears (Ursus americanus) at Lake Tahoe (CA). 2018 Ph.D., UCB
  6. *Branch, C., Elevation Related Variation in Climatic Harshness is Associated with Premating Mechanisms in Food-caching Mountain Chickadees (Poecile gambeli). 2018 Ph.D., UNR
  7. *Kozlovsky, D., The relationship between environment, behavior, cognition and the brain in specialized food-caching chickadees. 2018 Ph.D., UNR
  8. Wexler, Jeremy. Music Composition (Natural Reserve soundscapes). 2017, Masters, UCB
  9. Cassidy, Laura. The social and environmental turn in late 20th century art: a case study of Helen and Newton Harrison after modernism. 2017, Ph.D., Stanford
  10. Zemenick, A., The Influence of Flower Visitor Identity on Network Structure and Floral Microbe Communities. 2017 Ph.D., UCD
  11. Longley, P., Quantifying the Effects of Changing Snowpack Dynamics on Hydrologic Partitioning at Multiple Spatial Scales. 2017 Masters, UNR
  12. Lundahl, M., Place, Literacy Learning, and Civic Engagement: A Case Study of Adventure Risk Challenge. 2017 Ph.D., UNR
  13. Sapp, J., The behavioral ecology and intraspecific interactions of socially parasitic dulotic ants. 2017 Ph.D., UCSC.
  14. Sholtes, J., On the magnitude and frequency of sediment transport in rivers. 2016 Ph.D., Colorado State University.
  16. Kramer, N., Great River Wood Dynamics in Northern Canada. 2016 Ph.D., Colorado State University.
  17. Stephens, E., Critical and Aesthetic Research in Environmental Art. 2015 Ph.D., UCD.
  18. **Krimmel, E., MLIS E-Portfolio. 2014 Masters, San Jose State University.
  19. Harms, P. A., Distribution of Tritium in Precipitation and Surface Water in California. 2014 Masters, CSU-East Bay.
  20. Migliarese, N. L. , “That’s My Kind of Animal!” Designing and Assessing an Outdoor Science Education Program with Children’s Megafaunaphilia in Mind”. 2011 Ph.D., UCB.
  22. Theunissen Van Mannen, J., Using stable isotopes to assess longitudinal diet patterns of black bears (Ursas americanus) in Great Smoky Mountains National Park. 2011 Masters, UT-Knoxville.
  23. Haiman, A., Levels of Variation in Evening Grosbeak (Coccothraustes vespertinus) Calls and Morphology. 2011 Masters, UCD.
  24. Zane, M. Reaeration of Sagehen Creek Near Truckee, CA. 2010 Bachelor’s, UCSB
  25. Moeser, C. D., Development, Analysis and Use of a Distributed Wireless Sensor Network for Quantifying Spatial Trends of Snow Depth and Snow Water Equivalence Around Meteorological Stations With and Without Snow Sensing Equipment. 2010 Masters, UNR.
  26. Melody, A. D., Active faulting and quaternary paleohydrology of the Truckee fault zone north of Truckee, California. 2009 Masters, HSU.
  27. Moriarty, K. M., American Marten Distributions over a 28 Year Period: Relationships with Landscape Change in Sagehen Creek Experimental Forest, California, USA. 2009 Masters, OSU.
  28. Gorin, S., Transforming Thoughts into Words: Using Photography and Dialogue to Help Immigrant Students Write with Voice. 2009 Masters, UCD.
  29. Vaillant, N.M., Sagehen Experimental Forest Past, Present, and Future: An Evaluation of the Fireshed Assessment Process. 2008 Ph.D., UCB.
  30. Hooper, C., Bridging the gap: the union of personal and academic growth. ARC: Delivering literacy and personal growth for English Language Learners through outdoor education. 2008 Masters, Sierra Nevada College.
  31. Harssema, E., Examining the effectiveness of a standards-based, outdoor education summer program for English Language Learners. 2008 Masters, Sierra Nevada College.
  32. Richardson, W., Avian Use, Nest-site Selection, and Nesting Success in Sierra Nevada Aspen. 2007 Ph.D., UNR.
  33. Stead, J.E., Exploring Reintroduction of Lahontan Cutthroat Trout in a Headwater Stream. 2007 Masters, UCD.
  34. Ferree, E.D., Tail White and the Influence of Male Attractiveness on Maternal Investment in Dark-Eyed Juncos. 2007 Ph.D., UCSC.
  35. Sears, C.L., Assessing distribution, habitat suitability, and site occupancy of Great Gray Owls (Strix nebulosa) in California. 2006 Masters, UCD.
  36. Strathmann, K.S., Local and landscape influences on meadow butterfly communities in northeastern Sierra Nevada. 2005 Masters, SFSU.
  37. Soroka, D.E., Time budgets of the Willow Flycatcher in the Central Sierra Nevada in relation to nest success. 2003 Masters, UCD.
  38. Hatfield, R.G., Determinants of species richness and abundance of bumble bees, Bombus spp. (Hymenoptera: Apidae), in montane meadows of the Sierra Nevada, California. 2003 Masters, SFSU.
  39. *Bardsley, T., Investigations toward understanding the spatial representativeness of SNOTEL measurements of snow water equivalence. 2002 Masters, UNR.
  40. Rademacher, L.K., Groundwater flowpaths, chemical weathering rates, and climate change patterns as determined using multiple environmental tracers. 2002 Ph.D., UCSB.
  41. Kluse, J.S., Abiotic Gradients and Grazing Effects on the Interactions of Dominant Meadow Grasses, Deschampsia cespitosa a Native, and Poa pratensis ssp.pratensis a Non-Native. 2002 Masters, UCB.
  42. Elderd, B.D., Changing rivers and streams: the effects of anthropogenic alteration of riparian systems on Mimulus guttatus. 2002 Ph.D., UCSC.
  43. Merrill, A.G., Variation in the Structure & Nitrogen Dynamics of Mountain Riparian Zones. 2001 Ph.D., UCB.
  44. Cain, J.W., Nest Success of Yellow Warblers (Dendroica petechia)and Willow Flychatchers (Empidonax traillii) in relation to predator activity in montane meadows of the Central Sierra Nevada, CA. 2001 Ph.D., CSUS.
  45. Bombay, H., Scale Perspectives in Habitat Selection and Reproductive Success for Willow Flycatchers (Empidonax traillii) in the Central Sierra Nevada, California. 1999 Ph.D., CSUS.
  46. Campbell, E.A., Influence of Streamflow and Predators on Habitat Choice by Trout. 1998 Ph.D., UCD.
  47. Miller, III, D.G., The Manzanita Leaf-gall Aphid,Tamalia coweni (Cockerell) (Homoptera: Aphididae) as a Model System for Studies in Elementary Social Behavior, Sex Allocation and Life History Evolution. 1997 Ph.D., UCB.
  48. Ellsworth, S.D., Factors Influencing the Distribution and Abundance of Macroinvertebrates on Stones in Sagehen Creek, California. 1997 Ph.D., UCD.
  49. Williams, M.P., Inhibition of conifer regeneration by an herbaceous perennial, Wyethia mollis, in the eastern Sierra Nevada, California. 1995 Ph.D., U of W.
  50. Shivik, J.A., Factors Influencing Space Use and Activity of Sagehen Basin Coyotes. 1995 Ph.D., UCB.
  51. Woodard, E.L., Behavior, Activity Patterns and Foraging Strategies of Beaver (Castor canadensis) on Sagehen Creek, California. 1994 Ph.D., UCB.
  52. Kleinhouse, S., Ecophysiology of mutualism in a uniques stream symbiosis between a polymorphic cyanobacterium, Nostoc parmelioides, and the larvae of the midge Cricotopus nostocicola. 1994 Ph.D., UCD.
  53. Galloway, L.F., Evolution of Life-History Characters and Phenotypic Plasticity in Mimulus guttatus. 1994 Ph.D., UCD.
  54. Smith, J.I., Environmental influence on the ontogeny, allometry and behavior of the Song Sparrow (Melospiza melodia). 1993 Ph.D., UCB.
  55. Roden, J.S., The physiological benefit of leaf flutter in poplars. 1992 Ph.D., UCD.
  56. Ashman, T.-L.R., The Factors Governing the Maintenance of Gynodioecy in Sidalcia oregana ssp.spicata (Malvaceae). 1991 Ph.D., UCD.
  57. Reynolds, M.D., The Ecology of Spacing Behavior in the Yellow-billed Magpie (Pica nuttalli). 1990, Ph.D. UCB.
  58. Olson, M.D., Upstream changes in native fish abundance after reservoir impoundment in California streams of the Lahontan Basin. 1988 Masters, UCB.
  59. Baker, R.J., Response of trout to beaver dams in Sagehen Creek, California. 1988 Masters, UCB.
  60. Martin, S.K., The ecology of the Pine Marten (Martes americana) at Sagehen Creek, California. 1987 Ph.D., UCB.
  61. Cylinder, P., Competition for Pollination in a Montane Meadow. 1987 Ph.D., UCB
  62. Yoder-Williams, M., Quantitative effects of spider predation on arthropod populations in the eastern Sierra. 1986 Masters, U of Tn.
  63. Johnson, B.R., Influence of soil moisture, herbaceous competition, and small mammals on lodgepole pine (Pinus contorta var. murrayana [Grey & Balf] Engelm) seedling establishment in meadows of California. 1986 Masters, UCD.
  64. Berner, M. L., Sagehen Creek Field Station Planning and Site Design. 1985 Master of Landscape Architecture, UCB.
  65. Decker, L.M., Distribution, abundance and coexistence of two species of sucker (Catostomus) in Sagehen Creek, California, and their status in the western Lahontan Basin. 1984 Masters, UCB.
  66. Dedon, M.F., A test of wildlife habitat relationship models for Black Oaks and mixed conifer habitats in California. 1982 Ph.D., UCB.
  67. Zielinski, W.J., Food habits, activity patterns, and ectoparasites of the pine marten at Sagehen Creek, California. 1981 Ph.D., UCB.
  68. West, C.I., Management Options for the Sagehen Creek Basin, Tahoe National Forest, California. [contains extensive background information & metadata]. 1982 Masters of Forestry, UCB.
  69. Vondracek, B.C., The energetics of the Tahoe sucker, Catostomus tahoensis, in cyclic and constant temperatures. 1981 Masters, UCB.
  70. Spencer, W.D., Pine marten habitat preference at Sagehen Creek, California. 1981 Ph.D., UCB.
  71. Pitcher, D.C., Ecological Effects of Fire on Stand Structure and Fuel Dynamics in Red Fir Forests of Mineral King, Sequoia National Park, California. 1981 Ph.D., UCB.
  72. Cohen, Y., Ecological and morphological aspects of the breeding bird community of Blodgett Forest, California. 1981 Ph.D., UCB.
  73. Raphael, M.G., Utilization of dead standing trees by breeding birds at Sagehen Creek, California. 1980 Ph.D., UCB.
  74. Marrin, D.L., Food selectivity and habitat utilization by introduced trouts and non-game fishes in subalpine lakes. 1980 Masters, UCB.
  75. Busher, P.E., The population dynamics and behavior of beavers in the Sierra Nevada. 1980, Ph.D. UNR.
  76. Osaki, S.K., Impact of herbicide use on wildlife populations near Sagehen Creek. 1979, Masters UCB.
  77. Sharples, F.E., Resource partitioning in four sympatric species of Eutamias in the Northern Sierra Nevada. 1978, Masters UCD.
  78. Docekal, E.M., Home range, activities and diet of Belding’s ground squirrel (Spermophilus beldingi) at Sagehen Creek, California. 1978, Masters SFSU.
  79. Smith, S.F., Social structure of chipmunk populations and its relation to resource availability and exploitation. 1977, Ph.D. UCB.
  80. Schempf, P.F., Furbearers in the mountainous regions of California. 1977, Ph.D. UCB.
  81. Savidge, J.A., Effects on wildlife of herbicide-induced habitat change, Tahoe National Forest, California. 1977 Masters, UCB.
  82. Escherich, P.C., Social biology of the bushy-tailed woodrat. 1977, Ph.D. UCB.
  83. Bronson, M.T., Altitudinal variation in the annual cycle and life history of the golden-mantled ground squirrel (Spermophilus lateralis). 1977 Ph.D., UCB.
  84. Trowbridge, B.A., Plants of upper Sagehen Creek drainage basin, Nevada County, California. 1975, Masters SFSU.
  85. Busher, P.E., Movements and activities of beavers, Castor canadensis, on Sagehen Creek, California. 1975 Masters, SFSU.
  86. Beedy, E. C., Avifaunal complexity and forest structure in the Sierra Nevada of California. 1975, Masters UCD.
  87. Cameron, E.A., Bionomics and impact of the lodgepole terminal weevil, (Pissodes termialis Hopping (Coleoptera: Curculionidae), in the Sierra Nevada of California. 1974 Ph.D., UCB.
  88. Byrne, S., Dominance relationships and comparative ecology of two sympatric chipmunks (Eutamius). 1973, Masters UCB.
  89. Yost, W.F., Evaluation of Two Trout. 1972 Masters, UCB.
  90. White, J.M., Breeding biology and feeding patterns of the Oregon junco in two Sierra Nevada habitats. 1972, Ph.D. UCB.
  91. Reichart, H.A., Small mammal populations in six Sagehen Creek basin habitats. 1972, Masters UCB.
  92. Fitzgerald, B.M., The role of weasel predation in cyclic population changes of the montane vole (Microtus montanus). 1972, Ph.D. UCB.
  93. Beaver, D.L., Avian species diversity and habitat use in forests in the Sierra Nevada California. 1972, Ph.D. UCB.
  94. Balgooyen, T.G., Behavior and ecology of the American Kestrel (Falco sparverius). 1972, Ph.D. UCB.
  95. Taylor, D., Growth, decline and equilibrium in a beaver population in Sagehen Creek, California. 1970, Ph.D. UCB.
  96. Rae, S.P., Studies in the ecology of Mason Bog, Sagehen Creek, California. 1970, Ph.D. UCD.
  97. Hawthorne, V.M., Movements and food habits of coyotes in the Sagehen Creek basin and vicinity. 1970, Masters UNR.
  98. Bock, C.E., The ecology and behavior of the Lewis woodpecker (Asyndesmus lewis). 1970, Ph.D. UCB.
  99. Lea, R.N., Ecology of the Lahontan cutthroat trout, Salmo clarki henshawi, in Independence Lake, California. 1968, Masters UCB.
  100. Nee, J.A., The biology of the yellow-bellied marmot (Marmota flaviventris) in Sagehen Creek basin. 1967, Ph.D. UCB.
  101. Payen, L. A., Prehistoric rock art in the Northern Sierra Nevada, California. 1966 Masters, Sacramento State College.
  102. Evans, D.H., Life history of the Lahonton redside Richardsonius egregius (Girard), in Lake Tahoe. 1966 Masters, UCB.
  103. Wallis, J.R., A factor analysis of soil erosion and stream sediment in Northern California. 1965, Ph.D. UCB.
  104. Seegrist, D.W., The trout fishery of Sagehen Creek, Nevada County, California. 1965 Ph.D., UCB.
  105. Barnes, A.M., A revision of the genus Anomiopsyllus Baker, 1904 (Siphonaptera, Hystrichopsyllidae) with studies on the biology of Anomiopsyllus Falsicalifornicus. 1963, Ph.D. UCB.
  106. Roberts, D.R., Rodent movements in a cutover forest of the Sierra Nevada, California. 1962 Ph.D., UCB.
  107. Sherburne, F.C., Jr., Coexistence and Competition in Two Species of Legume Aphids. 1961 Ph.D., UCB.
  108. Behnke, R.J., Taxonomy of the cutthroat trout of the Great Basin with notes on the rainbow series. 1960 Masters, UCB.
  109. Seegrist, D.W., Age and growth of trout in Sagehen Creek, California. 1959, Masters UCB.
  110. Dietsch, E.L., The ecology and food habits of the sculpin Cottus beldingi in relation to the eastern brook trout, Salvelinus fontinalis. 1959, Masters UCB.
  111. Gard, R., The effects of beaver on trout in Sagehen Creek, California. 1958, Ph.D. UCB.
  112. Schnell, J.H., Nesting behavior and food habits of goshawks in the Sierra Nevada of California. 1957 Masters, UCB.
  113. Hoffmann, R.S., Population fluctuations of small mammals and sooty grouse in California. 1956, Masters UCB.
  114. Hall, J.G., Willow and aspen in the ecology of beaver on Sagehen Creek, California. 1956 Ph.D., UCB.
  115. Brock, E.M., Biology of the midge Cricotopus inhabiting the Alga Nostoc in California. 1956, Masters UCB.
  116. Kilgore, B.M., Breeding bird populations in managed and unmanaged stands of Sequoia giantea. 1954, Ph.D. UCB.
  117. Jones, A.C., Age, growth and reproduction of the sculpin (Cottus beldingi) in Sagehen Creek, Nevada-Sierra Counties, California. 1954, Masters UCB.
  118. Flittner, G.A., The composition and distribution of the fish populations in Sagehen Creek, Nevada-Sierra Counties, California. 1953, Ph.D. UCB.
  119. Newman, M.A., The social behavior of two species of juvenile salmonids. 1951, Masters UCB.


* UNR Ph.D.s (and some Masters) are awarded on the basis of a series of scientific publications in the field, not on a single thesis.

** San Jose State University Masters candidates may elect to submit a portfolio, rather than a thesis.